They don’t define “lifestyle” at all in that article. I imagine if I had to live in a cave and fight to get anything, I’d never reach whatever the natural max lifespan was, and would doubtless die in childbirth without skilled midwifery but it’s hard to imagine max being 38 when fertility for modern women lasts past 40, and we can live to 80 or 90. Are we not the same basic model?