There seems to be a lot of confusion regarding your question from users. If I understand it correctly, you’re wondering about when the human species became less of hunter-gatherers and more socially adept animals and when and why societal progress started suppressing aggressive behaviours in tribe-based expansion and control.

You referencing “protohuman traits” makes it a bit harder however since protohumans are before our own species of homo sapiens but I could imagine that a lot of those traits are closely related to early homo sapien society.

I don’t have a lot of answers for you but I think the answer maybe lies in sociological studies and things like Social Contract theory. Maybe it’s that slowly people got more rights to not be hurt by others because they could bring value to society as a whole in other ways and with the invention of agriculture, things like jobs became more viable and governments formed with laws then religion came along and what not too heavily influencing rights and wrongs.